The meaning of “תּוֹלָע”

“And the sons of Issachar; Tola, and Phuvah, and Job, and Shimron.”

“Of the sons of Issachar after their families: of Tola, the family of the Tolaites: of Pua, the family of the Punites:”

“And after Abimelech there arose to defend Israel Tola the son of Puah, the son of Dodo, a man of Issachar; and he dwelt in Shamir in mount Ephraim.”

“Now the sons of Issachar were, Tola, and Puah, Jashub, and Shimron, four.”

“And the sons of Tola; Uzzi, and Rephaiah, and Jeriel, and Jahmai, and Jibsam, and Shemuel, heads of their father’s house, to wit, of Tola: they were valiant men of might in their generations; whose number was in the days of David two and twenty thousand and six hundred.”